Vault-13

CODEXIS INC CDXS

COM · Common Stock · CUSIP 192005106

American protein engineering company

174 institutional holders
61,424,34161.4m shares reported held*
93.5% of all shares outstanding*
$372.5mreported value*

* Summed across managers. Shares that two managers' reports both contain – a subsidiary's holdings that its parent's report includes too, or a sub-adviser's in its client's – are counted once ($484.0k this quarter). Holder counts are more reliable. Shares outstanding: 65.7m as of 2022-11-01, from the company's own SEC filing; the percentage can still exceed 100%: lent-out shares are counted by the lender and the buyer, and some reports overlap without saying so. Implied quarter-end price ≈ $6.07. Options: 5 managers hold calls ($574.3k notional), 4 hold puts ($203.2k).

Biggest calls

Nantahala Capital Management, LLC — $303.0k50.0k shares
Jefferies Group LLC — $156.3k25.8k shares
SUSQUEHANNA INTERNATIONAL GROUP, LLP — $61.0k10.1k shares
SIMPLEX TRADING, LLC — $54.0k9.0k shares
CONCOURSE FINANCIAL GROUP SECURITIES, INC. — $00 shares

Biggest puts

Jefferies Group LLC — $121.2k20.0k shares
CITADEL ADVISORS LLC — $64.0k10.5k shares
SIMPLEX TRADING, LLC — $18.0k3.1k shares
CONCOURSE FINANCIAL GROUP SECURITIES, INC. — $00 shares

Options at notional value: the value of the shares the contracts control, as filers report them, not what the options cost. Not included in the holders, shares or buying above.

Major shareholders (over 5%)

Anyone who owns more than 5% of a class of the company's shares has to say so: on Schedule 13D if they may try to influence or control the company, on 13G if they're a passive investor, and update it as the stake changes. Each holder's latest filing, from 19 Dec 2024 on; percentages are of the share class, as the holder worked them out.

HolderStakeChangeShares Last filed
Opaleye Management Inc. 13G · passive investor no change · filed 2026-05-05 7.5% no change 6.8m≈ $15.3m 2026-05-05
Aberdeen Group plc 13G · institution -1.4 pts · filed 2026-07-20 7.5% -1.4 pts 6.8m≈ $15.3m 2026-07-20
Telemark Asset Management, LLC 13G · passive investor new · filed 2026-07-29 7.3% new 7.8m≈ $17.7m 2026-07-29
BlackRock, Inc. 13G · institution filed 2025-04-24 7.2% 6.0m≈ $13.5m 2025-04-24
LYTTON LAURENCE W 13G · passive investor +0.2 pts · filed 2026-08-13 5.2% +0.2 pts 5.6m≈ $12.6m 2026-08-13
INTEGRATED CORE STRATEGIES (US) LLC 13G · passive investor new · filed 2026-04-14 5.2% new 4.7m≈ $10.7m 2026-04-14

Fell to 5% or below in the last year: VANGUARD GROUP INC (0.0%, 2026-03-26)

Insider purchases and sales in Q3 2022

Shares bought or sold by the company's own officers, directors and 10% owners, in the open market or privately, as they reported to the SEC on Form 4 (due within two business days of the trade). Grants, option exercises, shares withheld for tax and gifts aren't included.

Officers and directors buying vs selling

Number of people, by quarter (10% owners not included)

BuyingSelling
404Q4 20: 0 buyingQ4 20: 0 sellingQ1 21: 0 buyingQ1 21: 0 sellingQ1 21Q2 21: 0 buyingQ2 21: 0 sellingQ3 21: 0 buyingQ3 21: 0 sellingQ3 21Q4 21: 0 buyingQ4 21: 4 sellingQ1 22: 0 buyingQ1 22: 3 sellingQ1 22Q2 22: 0 buyingQ2 22: 1 sellingQ3 22: 0 buyingQ3 22: 1 sellingQ3 22
Numbers
QuarterBuyingBoughtSellingSoldSold under a plan
Q4 20200—0——
Q1 20210—0——
Q2 20210—0——
Q3 20210—0——
Q4 20210—4$4.0m—
Q1 20220—3$134.3k—
Q2 20220—1$793.9k—
Q3 20220—1$674.3k—
DateInsiderTradeSharesPrice ValueTheir holding
2022-08-15 NICOLS JOHN J Director Sold 39,000 $8.78 $342.4k39.0k shares -4%
2022-08-12 NICOLS JOHN J Director Sold 38,500 $8.62 $331.9k38.5k shares -4%

Their holding: how much the trade added to or took from the insider's holding (direct, or the trust or fund named). The same trade is sometimes reported by more than one insider, such as a fund and the director who represents it; totals count it once. A value is left blank where the reported price is far from the market price (usually a typo).

Holders, Q3 2022

#ManagerLocationShares Value% of their portfolioChangeShares change
1 BAILLIE GIFFORD & CO 0.04% of their portfolio EDINBURGH, X0 7,068,1197.1m $42.8m $42.8m 0.04% —
2 BlackRock Inc. 0.00% of their portfolio New York, NY 5,481,5935.5m $33.2m $33.2m 0.00% —
3 Nantahala Capital Management, LLC 2.68% of their portfolio NEW CANAAN, CT 4,504,0094.5m $27.3m $27.3m 2.68% —
4 ARK Investment Management LLC 0.18% of their portfolio ST. PETERSBURG, FL 4,273,4854.3m $25.9m $25.9m 0.18% —
5 VANGUARD GROUP INC 0.00% of their portfolio Valley Forge, PA 3,515,4103.5m $21.3m $21.3m 0.00% —
6 Telemark Asset Management, LLC 2.57% of their portfolio BOSTON, MA 2,704,5002.7m $16.4m $16.4m 2.57% —
7 AMERIPRISE FINANCIAL INC 0.00% of their portfolio MINNEAPOLIS, MN 1,960,8882.0m $11.9m $11.9m 0.00% —
8 FMR LLC 0.00% of their portfolio BOSTON, MA 1,826,3571.8m $11.1m $11.1m 0.00% —
9 WELLINGTON MANAGEMENT GROUP LLP 0.00% of their portfolio Boston, MA 1,554,0691.6m $9.4m $9.4m 0.00% —
10 GILDER GAGNON HOWE & CO LLC 0.15% of their portfolio New York, NY 1,459,9131.5m $8.8m $8.8m 0.15% —
11 Nuveen Asset Management, LLC 0.00% of their portfolio CHICAGO, IL 1,406,7771.4m $8.5m $8.5m 0.00% —
12 JACOBS LEVY EQUITY MANAGEMENT, INC 0.06% of their portfolio FLORHAM PARK, NJ 1,327,6681.3m $8.0m $8.0m 0.06% —
13 STATE STREET CORP 0.00% of their portfolio BOSTON, MA 1,307,7451.3m $7.9m $7.9m 0.00% —
14 GOLDMAN SACHS GROUP INC 0.00% of their portfolio NEW YORK, NY 1,289,1291.3m $7.8m $7.8m 0.00% —
15 Opaleye Management Inc. 2.89% of their portfolio BOSTON, MA 1,155,0001.2m $7.0m $7.0m 2.89% —
16 Sumitomo Mitsui Trust Group, Inc. 0.01% of their portfolio CHIYODA-KU, TOKYO, M0 1,132,1241.1m $6.9m $6.9m 0.01% —
17 Amova Asset Management Americas, Inc. 0.09% of their portfolio New York, NY 1,132,1241.1m $6.9m $6.9m 0.09% —
18 GEODE CAPITAL MANAGEMENT, LLC 0.00% of their portfolio BOSTON, MA 1,124,5731.1m $6.8m $6.8m 0.00% —
19 WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC 0.03% of their portfolio CHICAGO, IL 1,090,4121.1m $6.6m $6.6m 0.03% —
20 CITADEL ADVISORS LLC 0.01% of their portfolio MIAMI, FL 787,769787.8k $4.8m $4.8m 0.01% —
21 TWO SIGMA ADVISERS, LP 0.01% of their portfolio NEW YORK, NY 768,600768.6k $4.7m $4.7m 0.01% —
22 KORNITZER CAPITAL MANAGEMENT INC /KS 0.10% of their portfolio SHAWNEE MISSION, KS 751,264751.3k $4.6m $4.6m 0.10% —
23 TWO SIGMA INVESTMENTS, LP 0.01% of their portfolio NEW YORK, NY 697,384697.4k $4.2m $4.2m 0.01% —
24 MORGAN STANLEY 0.00% of their portfolio NEW YORK, NY 622,033622.0k $3.8m $3.8m 0.00% —
25 CHARLES SCHWAB INVESTMENT MANAGEMENT INC 0.00% of their portfolio SAN FRANCISCO, CA 612,668612.7k $3.7m $3.7m 0.00% —
26 NORTHERN TRUST CORP 0.00% of their portfolio CHICAGO, IL 612,148612.1k $3.7m $3.7m 0.00% —
27 Allspring Global Investments Holdings, LLC 0.01% of their portfolio CHARLOTTE, NC 587,089587.1k $3.6m $3.6m 0.01% —
28 Prescott Group Capital Management, L.L.C. 1.00% of their portfolio Tulsa, OK 532,653532.7k $3.2m $3.2m 1.00% —
29 D. E. Shaw & Co., Inc. 0.01% of their portfolio New York, NY 525,334525.3k $3.2m $3.2m 0.01% —
30 Assenagon Asset Management S.A. 0.01% of their portfolio SENNINGERBERG, N4 492,682492.7k $3.0m $3.0m 0.01% —
31 Vivo Capital, LLC 0.26% of their portfolio Palo Alto, CA 388,144388.1k $2.4m $2.4m 0.26% —
32 Cubist Systematic Strategies, LLC 0.02% of their portfolio STAMFORD, CT 361,401361.4k $2.2m $2.2m 0.02% —
33 Walleye Capital LLC 0.05% of their portfolio NEW YORK, NY 327,541327.5k $2.0m $2.0m 0.05% —
34 MARSHALL WACE, LLP 0.00% of their portfolio LONDON, X0 309,633309.6k $1.9m $1.9m 0.00% —
35 ALGERT GLOBAL LLC 0.11% of their portfolio SAN FRANCISCO, CA 293,069293.1k $1.8m $1.8m 0.11% —
36 VOLORIDGE INVESTMENT MANAGEMENT, LLC 0.01% of their portfolio JUPITER, FL 292,453292.5k $1.8m $1.8m 0.01% —
37 BANK OF AMERICA CORP /DE/ 0.00% of their portfolio CHARLOTTE, NC 290,947290.9k $1.8m $1.8m 0.00% —
38 Legal & General Group Plc 0.00% of their portfolio London, X0 243,811243.8k $1.5m $1.5m 0.00% —
39 Bank of New York Mellon Corp 0.00% of their portfolio NEW YORK, NY 232,311232.3k $1.4m $1.4m 0.00% —
40 DIMENSIONAL FUND ADVISORS LP 0.00% of their portfolio AUSTIN, TX 229,252229.3k $1.4m $1.4m 0.00% —
41 BANK OF MONTREAL /CAN/ 0.00% of their portfolio TORONTO, A6 227,309227.3k $1.7m $1.7m 0.00% —
42 Knott David M Jr 0.58% of their portfolio SYOSSET, NY 222,107222.1k $1.3m $1.3m 0.58% —
43 ESSEX INVESTMENT MANAGEMENT CO LLC 0.21% of their portfolio BOSTON, MA 217,806217.8k $1.3m $1.3m 0.21% —
44 Squarepoint Ops LLC 0.01% of their portfolio NEW YORK, NY 201,802201.8k $1.2m $1.2m 0.01% —
45 JACOB ASSET MANAGEMENT OF NEW YORK LLC 1.24% of their portfolio HERMOSA BEACH, CA 188,513188.5k $1.1m $1.1m 1.24% —
46 EXCHANGE TRADED CONCEPTS, LLC 0.04% of their portfolio Oklahoma City, OK 187,472187.5k $1.1m $1.1m 0.04% —
47 MILLENNIUM MANAGEMENT LLC 0.00% of their portfolio NEW YORK, NY 170,842170.8k $1.0m $1.0m 0.00% —
48 Northwest & Ethical Investments L.P. 0.04% of their portfolio TORONTO, A6 169,409169.4k $1.0m $1.0m 0.04% —
49 PUTNAM INVESTMENTS LLC 0.00% of their portfolio BOSTON, MA 160,756160.8k $975.0k $975.0k 0.00% —
50 RENAISSANCE TECHNOLOGIES LLC 0.00% of their portfolio NEW YORK, NY 154,992155.0k $939.0k $939.0k 0.00% —
51 Dynamic Technology Lab Private Ltd 0.09% of their portfolio SINGAPORE, U0 152,127152.1k $922.0k $922.0k 0.09% —
52 EATON VANCE MANAGEMENT 0.00% of their portfolio BOSTON, MA 150,340150.3k $911.0k $911.0k 0.00% —
53 AQR CAPITAL MANAGEMENT LLC 0.00% of their portfolio Greenwich, CT 146,298146.3k $887.0k $887.0k 0.00% —
54 SILVERARC CAPITAL MANAGEMENT, LLC 0.37% of their portfolio BOSTON, MA 145,437145.4k $881.0k $881.0k 0.37% —
55 WHITE PINE CAPITAL LLC 0.36% of their portfolio BLOOMINGTON, MN 143,246143.2k $868.0k $868.0k 0.36% —
56 Candriam S.C.A. 0.01% of their portfolio STRASSEN, N4 142,953143.0k $866.0k $866.0k 0.01% —
57 GSA CAPITAL PARTNERS LLP 0.11% of their portfolio LONDON, X0 142,410142.4k $863.0k $863.0k 0.11% —
58 Swiss National Bank 0.00% of their portfolio ZURICH, V8 138,000138.0k $836.0k $836.0k 0.00% —
59 Schonfeld Strategic Advisors LLC 0.01% of their portfolio NEW YORK, NY 120,200120.2k $728.0k $728.0k 0.01% —
60 Engineers Gate Manager LP 0.04% of their portfolio NEW YORK, NY 116,223116.2k $704.0k $704.0k 0.04% —
61 OCCUDO QUANTITATIVE STRATEGIES LP 0.05% of their portfolio FORT LAUDERDALE, FL 107,532107.5k $652.0k $652.0k 0.05% —
62 Parkman Healthcare Partners LLC 0.18% of their portfolio STAMFORD, CT 105,319105.3k $638.0k $638.0k 0.18% —
63 BNP PARIBAS FINANCIAL MARKETS 0.00% of their portfolio PARIS, I0 102,471102.5k $621.0k $621.0k 0.00% —
64 Russell Investments Group, Ltd. 0.00% of their portfolio Seattle, WA 98,37998.4k $595.0k $595.0k 0.00% —
65 ExodusPoint Capital Management, LP 0.01% of their portfolio NEW YORK, NY 92,81492.8k $562.0k $562.0k 0.01% —
66 Private Advisor Group, LLC 0.01% of their portfolio MORRISTOWN, NJ 87,05987.1k $528.0k $528.0k 0.01% —
67 CREDIT SUISSE AG/ 0.00% of their portfolio ZURICH, V8 80,76080.8k $490.0k $490.0k 0.00% —
68 RHUMBLINE ADVISERS 0.00% of their portfolio BOSTON, MA 79,20879.2k $480.0k $480.0k 0.00% —
69 UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC 0.00% of their portfolio NEW YORK, NY 78,94078.9k $478.4k $478.4k 0.00% —
70 BRADLEY FOSTER & SARGENT INC/CT 0.01% of their portfolio HARTFORD, CT 76,48276.5k $463.0k $463.0k 0.01% —
71 CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM 0.00% of their portfolio WEST SACRAMENTO, CA 74,73874.7k $453.0k $453.0k 0.00% —
72 AIA Group Ltd 0.03% of their portfolio HONG KONG, K3 72,19572.2k $438.0k $438.0k 0.03% —
73 AlphaCrest Capital Management LLC 0.03% of their portfolio New York, NY 70,96571.0k $430.0k $430.0k 0.03% —
74 Conestoga Capital Advisors, LLC 0.01% of their portfolio WAYNE, PA 70,91170.9k $430.0k $430.0k 0.01% —
75 Tekla Capital Management LLC 0.02% of their portfolio Boston, MA 69,12569.1k $419.0k $419.0k 0.02% —
76 ALLIANCEBERNSTEIN L.P. 0.00% of their portfolio NASHVILLE, TN 61,57061.6k $373.0k $373.0k 0.00% —
77 Jump Financial, LLC 0.01% of their portfolio Chicago, IL 61,00061.0k $370.0k $370.0k 0.01% —
78 LORD, ABBETT & CO. LLC 0.00% of their portfolio JERSEY CITY, NJ 60,72360.7k $368.0k $368.0k 0.00% —
79 JACKSON SQUARE CAPITAL, LLC 0.18% of their portfolio SAN FRANCISCO, CA 59,89759.9k $363.0k $363.0k 0.18% —
80 JPMORGAN CHASE & CO 0.00% of their portfolio NEW YORK, NY 58,93858.9k $357.0k $357.0k 0.00% —
81 ROYAL BANK OF CANADA 0.00% of their portfolio TORONTO, A6 49,37349.4k $299.0k $299.0k 0.00% —
82 SG Americas Securities, LLC 0.00% of their portfolio NEW YORK, NY 43,39743.4k $263.0k $263.0k 0.00% —
83 PRICE T ROWE ASSOCIATES INC /MD/ 0.00% of their portfolio BALTIMORE, MD 42,71442.7k $259.0k $259.0k 0.00% —
84 BLAIR WILLIAM & CO/IL 0.00% of their portfolio CHICAGO, IL 42,66842.7k $259.0k $259.0k 0.00% —
85 NEW YORK STATE COMMON RETIREMENT FUND 0.00% of their portfolio ALBANY, NY 38,61038.6k $234.0k $234.0k 0.00% —
86 STRS OHIO 0.00% of their portfolio COLUMBUS, OH 36,80036.8k $223.0k $223.0k 0.00% —
87 Qube Research & Technologies Ltd 0.00% of their portfolio LONDON, X0 35,06835.1k $213.0k $213.0k 0.00% —
88 Atom Investors LP 0.07% of their portfolio AUSTIN, TX 34,37034.4k $208.0k $208.0k 0.07% —
89 Campbell & CO Investment Adviser LLC 0.05% of their portfolio BALTIMORE, MD 33,72033.7k $204.0k $204.0k 0.05% —
90 Inspire Investing, LLC 0.03% of their portfolio MERIDIAN, ID 33,28833.3k $202.0k $202.0k 0.03% —
91 MetLife Investment Management, LLC 0.00% of their portfolio Whippany, NJ 32,96333.0k $199.8k $199.8k 0.00% —
92 Verition Fund Management LLC 0.00% of their portfolio GREENWICH, CT 31,82031.8k $192.0k $192.0k 0.00% —
93 Invesco Ltd. 0.00% of their portfolio ATLANTA, GA 29,29029.3k $178.0k $178.0k 0.00% —
94 AMERICAN INTERNATIONAL GROUP, INC. 0.00% of their portfolio New York, NY 29,06229.1k $176.0k $176.0k 0.00% —
95 XTX Topco Ltd 0.04% of their portfolio LONDON, X0 28,74128.7k $174.0k $174.0k 0.04% —
96 STATE BOARD OF ADMINISTRATION OF FLORIDA RETIREMENT SYSTEM 0.00% of their portfolio TALLAHASSEE, FL 28,50028.5k $173.0k $173.0k 0.00% —
97 Versor Investments LP 0.01% of their portfolio New York, NY 27,30027.3k $165.0k $165.0k 0.01% —
98 DEUTSCHE BANK AG\ 0.00% of their portfolio FRANKFURT AM MAIN, 2M 26,73726.7k $162.0k $162.0k 0.00% —
99 PANAGORA ASSET MANAGEMENT INC 0.00% of their portfolio BOSTON, MA 26,65626.7k $162.0k $162.0k 0.00% —
100 Mirae Asset Global Investments Co., Ltd. 0.00% of their portfolio SEOUL, M5 26,61326.6k $161.0k $161.0k 0.00% —